Updated: July 8, 2025

The peppered moth (Biston betularia) is one of the most famous examples in evolutionary biology, largely due to its striking color variations and the way these variations have been studied to understand natural selection. This unassuming insect has played a pivotal role in illustrating how species adapt to their environments through changes in appearance. In this article, we will explore the quick facts about the peppered moth’s color variations, their significance, causes, and impact on evolutionary science.

Overview of the Peppered Moth

The peppered moth is a species native to Europe and North America. It is a nocturnal moth belonging to the family Geometridae. The name “peppered” comes from its speckled wing patterns that help it camouflage against tree bark and lichens. The moth’s primary predators are birds, which hunt visually during the day, making camouflage a vital adaptation for survival.

The Two Main Color Variations

Peppered moths are best known for their two main color morphs:

  • Typica (Light-colored morph): This variant has light wings mottled with black or dark gray speckles, resembling the appearance of lichen-covered tree bark.
  • Carbonaria (Dark-colored morph): This variant is predominantly black or very dark gray, almost completely obscuring the speckled pattern.

These two forms coexist but vary dramatically in frequency depending on environmental conditions.

Historical Significance: Industrial Melanism

The peppered moth’s color variations became a landmark example of industrial melanism. During the Industrial Revolution in England (18th to 19th centuries), pollution from factories darkened tree bark with soot and killed off light-colored lichens. This environmental change caused a shift in selective pressures:

  • The light-colored typica moths became more visible against soot-darkened trees and were thus more likely to be eaten by birds.
  • The dark-colored carbonaria moths were better camouflaged on sooty bark and survived at higher rates.

As a result, the population shifted dramatically toward the carbonaria form in industrial areas—a textbook example of natural selection driving evolutionary change.

Genetics Behind Color Variations

The differences between typica and carbonaria are controlled primarily by genetic factors. Research identifies that a single gene locus with multiple alleles influences the coloration pattern. The carbonaria morph results from a dominant allele causing increased melanin production, leading to darker pigmentation.

Recent genomic studies have pinpointed that this mutation involves a transposable element insertion within the gene cortex. This gene plays an essential role in butterfly and moth wing patterning, affecting pigment deposition during development.

Beyond Typica and Carbonaria: Intermediate Forms

In addition to the two classic forms, there are intermediate or rare morphs exhibiting varying degrees of darkness or speckling. These include:

  • Insularia: A lighter form with reduced speckling.
  • Punctata: Characterized by distinct black dots against a lighter background.
  • Various other localized variants influenced by geographic and environmental factors.

These variants demonstrate that peppered moth coloration exists on a continuum rather than merely as two discrete states.

The Role of Camouflage and Predation

Camouflage is critical for peppered moth survival due to their susceptibility to bird predation during daylight hours when resting on tree trunks. Studies using predation experiments have demonstrated that birds preferentially prey on individuals whose coloration contrasts with their resting background.

This dynamic creates strong selective pressure favoring moths whose wing patterns best match their immediate environment, leading to fluctuations in morph frequencies corresponding to changes in habitat coloration.

Environmental Influence Across Regions

The prevalence of different color morphs varies geographically based on factors such as:

  • Pollution levels: Industrial regions showcase higher carbonaria frequencies.
  • Lichen abundance: Lichens provide background matching for typica forms.
  • Forest type and tree species: Different bark textures and colors influence which morph is favored.
  • Urbanization: Modern clean air acts have led to reductions in soot deposits, causing a resurgence of typica forms in many areas.

These environmental factors continually shape local populations through natural selection.

Evidence Supporting Natural Selection

The peppered moth story has been scrutinized over decades, providing extensive evidence for natural selection:

  • Mark-release-recapture experiments: Demonstrated differential predation rates favoring cryptic morphs.
  • Historical records: Correlated shifts in morph frequency with pollution data.
  • Genetic analyses: Confirmed inheritance patterns consistent with selective pressures.
  • Reversal trends: Populations reverting toward typica following pollution reduction strongly support adaptive responses rather than random change.

While some early studies faced criticism regarding methodology or interpretation, subsequent rigorous research has reaffirmed the peppered moth as a robust example of evolution in action.

Importance in Evolutionary Biology Education

The peppered moth remains one of the most accessible case studies for demonstrating evolution by natural selection due to its visible phenotypic variation and well-documented historical context. It helps illustrate key concepts such as:

  • Adaptation
  • Mutation
  • Genetic dominance
  • Environmental influence
  • Selective pressure
  • Evolutionary dynamics over short timescales

This case continues to inspire scientific inquiry and public understanding about how organisms evolve in response to environmental changes.

Current Research Directions

Modern research on peppered moths focuses on areas such as:

  • Detailed genomic mapping of pigmentation genes.
  • Molecular mechanisms underlying melanin regulation.
  • Expanded ecological studies examining urban environments.
  • Climate change impacts on distribution and coloration patterns.
  • Comparative analyses with other melanistic species.

These efforts aim to deepen our understanding of evolutionary mechanisms at genetic, developmental, and ecological levels.

Summary: What Makes Peppered Moth Color Variations Fascinating?

The peppered moth’s color variations provide a compelling window into evolution through natural selection because they:

  1. Exhibit clear, genetically based polymorphism that affects survival.
  2. Respond dynamically to environmental changes driven by human activity.
  3. Demonstrate rapid evolutionary adaptation observable within human lifetimes.
  4. Serve as a model system bridging genetics, ecology, and evolution.
  5. Highlight how seemingly small organisms reveal big truths about life’s diversity and resilience.

Understanding these variations not only enriches our knowledge of biology but also emphasizes the profound impact humans have on ecosystems—and how species adapt accordingly.


Whether you are an aspiring biologist or simply curious about nature’s intricate designs, the story of the peppered moth’s color variations is an elegant testament to evolution unfolding right before our eyes.

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